diff --git a/src/material/button/testing/button-harness.spec.ts b/src/material/button/testing/button-harness.spec.ts
index 7f3ccda51ef8..496add10ba91 100644
--- a/src/material/button/testing/button-harness.spec.ts
+++ b/src/material/button/testing/button-harness.spec.ts
@@ -1,4 +1,4 @@
-import {Component} from '@angular/core';
+import {Component, signal} from '@angular/core';
 import {ComponentFixture, TestBed} from '@angular/core/testing';
 import {Platform} from '@angular/cdk/platform';
 import {HarnessLoader, parallel} from '@angular/cdk/testing';
@@ -7,22 +7,21 @@ import {MatButtonModule} from '../module';
 import {MatIconModule} from '../../icon';
 import {MatIconHarness} from '../../icon/testing';
 import {MatButtonHarness} from './button-harness';
+import {MatButtonAppearance} from '../button-base';
 
 describe('MatButtonHarness', () => {
   let fixture: ComponentFixture<ButtonHarnessTest>;
   let loader: HarnessLoader;
-  let platform: Platform;
 
   beforeEach(() => {
     fixture = TestBed.createComponent(ButtonHarnessTest);
     fixture.detectChanges();
     loader = TestbedHarnessEnvironment.loader(fixture);
-    platform = TestBed.inject(Platform);
   });
 
   it('should load all button harnesses', async () => {
     const buttons = await loader.getAllHarnesses(MatButtonHarness);
-    expect(buttons.length).toBe(16);
+    expect(buttons.length).toBe(17);
   });
 
   it('should load button with exact text', async () => {
@@ -41,7 +40,7 @@ describe('MatButtonHarness', () => {
   it('should filter by whether a button is disabled', async () => {
     const enabledButtons = await loader.getAllHarnesses(MatButtonHarness.with({disabled: false}));
     const disabledButtons = await loader.getAllHarnesses(MatButtonHarness.with({disabled: true}));
-    expect(enabledButtons.length).toBe(14);
+    expect(enabledButtons.length).toBe(15);
     expect(disabledButtons.length).toBe(2);
   });
 
@@ -79,7 +78,7 @@ describe('MatButtonHarness', () => {
     const button = await loader.getHarness(MatButtonHarness.with({text: 'Basic button'}));
     await button.click();
 
-    expect(fixture.componentInstance.clicked).toBe(true);
+    expect(fixture.componentInstance.clicked()).toBe(true);
   });
 
   it('should not click a disabled button', async () => {
@@ -88,14 +87,14 @@ describe('MatButtonHarness', () => {
     // cancel dispatched click events on disabled buttons. We skip this check on Edge and Firefox.
     // See: https://bugzilla.mozilla.org/show_bug.cgi?id=1582570 and:
     // https://stackoverflow.com/questions/32377026/disabled-button-is-clickable-on-edge-browser
-    if (platform.FIREFOX) {
+    if (TestBed.inject(Platform).FIREFOX) {
       return;
     }
 
     const button = await loader.getHarness(MatButtonHarness.with({text: 'Filled button'}));
     await button.click();
 
-    expect(fixture.componentInstance.clicked).toBe(false);
+    expect(fixture.componentInstance.clicked()).toBe(false);
   });
 
   it('should be able to handle nested harnesses', async () => {
@@ -127,6 +126,7 @@ describe('MatButtonHarness', () => {
       'basic',
       'basic',
       'basic',
+      'basic',
       'icon',
       'fab',
       'mini-fab',
@@ -151,6 +151,7 @@ describe('MatButtonHarness', () => {
       'filled',
       'elevated',
       'outlined',
+      'tonal',
       null,
       null,
       null,
@@ -166,16 +167,25 @@ describe('MatButtonHarness', () => {
     const button = await loader.getHarness(MatButtonHarness.with({appearance: 'filled'}));
     expect(await button.getText()).toBe('Filled button');
   });
+
+  it('should get the appearance of a button with a dynamic appearance', async () => {
+    const button = await loader.getHarness(
+      MatButtonHarness.with({selector: '#dynamic-appearance'}),
+    );
+    expect(await button.getAppearance()).toBe('tonal');
+    fixture.componentInstance.dynamicAppearance.set('filled');
+    expect(await button.getAppearance()).toBe('filled');
+  });
 });
 
 @Component({
   // Include one of each type of button selector to ensure that they're all captured by
   // the harness's selector.
   template: `
-    <button id="basic" type="button" matButton (click)="clicked = true">
+    <button id="basic" type="button" matButton (click)="clicked.set(true)">
       Basic button
     </button>
-    <button id="flat" type="button" matButton="filled" disabled (click)="clicked = true">
+    <button id="flat" type="button" matButton="filled" disabled (click)="clicked.set(true)">
       Filled button
     </button>
     <button id="raised" type="button" matButton="elevated">Elevated button</button>
@@ -194,6 +204,7 @@ describe('MatButtonHarness', () => {
     <a id="anchor-flat" matButton="filled">Filled anchor</a>
     <a id="anchor-raised" matButton="elevated" disabled>Elevated anchor</a>
     <a id="anchor-stroked" matButton="outlined">Stroked anchor</a>
+    <a id="dynamic-appearance" [matButton]="dynamicAppearance()">Stroked anchor</a>
     <a id="anchor-icon" matIconButton>Icon anchor</a>
     <a id="anchor-fab" matFab>Fab anchor</a>
     <a id="anchor-mini-fab" matMiniFab>Mini Fab anchor</a>
@@ -201,6 +212,6 @@ describe('MatButtonHarness', () => {
   imports: [MatButtonModule, MatIconModule],
 })
 class ButtonHarnessTest {
-  disabled = true;
-  clicked = false;
+  clicked = signal(false);
+  dynamicAppearance = signal<MatButtonAppearance>('tonal');
 }
diff --git a/src/material/button/testing/button-harness.ts b/src/material/button/testing/button-harness.ts
index e99545d493bb..65bc7fc0040f 100644
--- a/src/material/button/testing/button-harness.ts
+++ b/src/material/button/testing/button-harness.ts
@@ -16,10 +16,14 @@ import {ButtonAppearance, ButtonHarnessFilters, ButtonVariant} from './button-ha
 
 /** Harness for interacting with a mat-button in tests. */
 export class MatButtonHarness extends ContentContainerComponentHarness {
-  // TODO(jelbourn) use a single class, like `.mat-button-base`
-  static hostSelector = `[matButton], [mat-button], [matIconButton], [matFab], [matMiniFab],
-    [mat-raised-button], [mat-flat-button], [mat-icon-button], [mat-stroked-button], [mat-fab],
-    [mat-mini-fab]`;
+  // Note: `.mat-mdc-button-base` should be enough for all buttons, however some apps are using
+  // the harness without actually having an applied button. Keep the attributes for backwards
+  // compatibility.
+
+  /** Selector for the harness. */
+  static hostSelector = `.mat-mdc-button-base, [matButton], [mat-button], [matIconButton],
+    [matFab], [matMiniFab], [mat-raised-button], [mat-flat-button], [mat-icon-button],
+    [mat-stroked-button], [mat-fab], [mat-mini-fab]`;
 
   /**
    * Gets a `HarnessPredicate` that can be used to search for a button with specific attributes.